How to format your references using the ZooKeys citation style

This is a short guide how to format citations and the bibliography in a manuscript for ZooKeys. For a complete guide how to prepare your manuscript refer to the journal's instructions to authors.

Using reference management software

Typically you don't format your citations and bibliography by hand. The easiest way is to use a reference manager:

PaperpileThe citation style is built in and you can choose it in Settings > Citation Style or Paperpile > Citation Style in Google Docs.
EndNoteFind the style here: output styles overview
Mendeley, Zotero, Papers, and othersThe style is either built in or you can download a CSL file that is supported by most references management programs.
BibTeXBibTeX syles are usually part of a LaTeX template. Check the instructions to authors if the publisher offers a LaTeX template for this journal.

Journal articles

Those examples are references to articles in scholarly journals and how they are supposed to appear in your bibliography.

Not all journals organize their published articles in volumes and issues, so these fields are optional. Some electronic journals do not provide a page range, but instead list an article identifier. In a case like this it's safe to use the article identifier instead of the page range.

A journal article with 1 author
Kreeger K (2003) The learning curve. Nature 424: 234–235.
A journal article with 2 authors
Gitlin AD, Nussenzweig MC (2015) Immunology: Fifty years of B lymphocytes. Nature 517: 139–141.
A journal article with 3 authors
Biswas AK, Atulasimha J, Bandyopadhyay S (2014) An error-resilient non-volatile magneto-elastic universal logic gate with ultralow energy-delay product. Scientific reports 4: 7553.
A journal article with 4 or more authors
Young ED, Simon JI, Galy A, Russell SS, Tonui E, Lovera O (2005) Supra-canonical 26Al/27Al and the residence time of CAIs in the solar protoplanetary disk. Science (New York, N.Y.) 308: 223–227.

Books and book chapters

Here are examples of references for authored and edited books as well as book chapters.

An authored book
Cherniakov M (2003) An Introduction to Parametric Digital Filters and Oscillators. John Wiley & Sons, Ltd, Chichester, UK.
An edited book
Bressoud D (2016) Teaching and Learning of Calculus. Ghedamsi I, Martinez-Luaces V, Törner G (Eds). Springer International Publishing, Cham, VII, 37 p pp.
A chapter in an edited book
Croft J (2007) Creative Use of PNG Transparency in Web Design. In: Boulton M, Clarke A, Collison S, Croft J, Featherstone D, Lloyd I, Marcotte E, Rubin D, Weychert R (Eds), Web Standards Creativity: Innovations in Web Design with XHTML, CSS, and DOM Scripting. Apress, Berkeley, CA, 108–128.

Web sites

Sometimes references to web sites should appear directly in the text rather than in the bibliography. Refer to the Instructions to authors for ZooKeys.

Blog post
Fang J (2015) Bonobo Tool Use Resembles that of Our Early Stone Age Ancestors. IFLScience. Available from: (October 30, 2018).


This example shows the general structure used for government reports, technical reports, and scientific reports. If you can't locate the report number then it might be better to cite the report as a book. For reports it is usually not individual people that are credited as authors, but a governmental department or agency like "U. S. Food and Drug Administration" or "National Cancer Institute".

Government report
Government Accountability Office (2004) Telecommunications: Intelsat Privatization and the Implementation of the ORBIT Act. U.S. Government Printing Office, Washington, DC

Theses and dissertations

Theses including Ph.D. dissertations, Master's theses or Bachelor theses follow the basic format outlined below.

Doctoral dissertation
Bigler WH (2010) Increased substance abuse treatment for adolescents. Doctoral dissertation. California State University, Long Beach

News paper articles

Unlike scholarly journals, news papers do not usually have a volume and issue number. Instead, the full date and page number is required for a correct reference.

New York Times article
Sophia Kishkovsky; Compiled by (2006) Arts, Briefly; 2 Missing Russian Works Recovered. New York Times: B8.

In-text citations

References should be cited in the text by name and year in parentheses:

This sentence cites one reference (Kreeger 2003).
This sentence cites two references (Kreeger 2003, Gitlin and Nussenzweig 2015).

Here are examples of in-text citations with multiple authors:

  • Two authors: (Gitlin and Nussenzweig 2015)
  • Three or more authors: (Young et al. 2005)

About the journal

Full journal titleZooKeys
ISSN (print)1313-2989
ISSN (online)1313-2970
ScopeAnimal Science and Zoology
Ecology, Evolution, Behavior and Systematics

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