How to format your references using the ZooKeys citation style

This is a short guide how to format citations and the bibliography in a manuscript for ZooKeys. For a complete guide how to prepare your manuscript refer to the journal's instructions to authors.

Using reference management software

Typically you don't format your citations and bibliography by hand. The easiest way is to use a reference manager:

PaperpileThe citation style is built in and you can choose it in Settings > Citation Style or Paperpile > Citation Style in Google Docs.
EndNoteFind the style here: output styles overview
Mendeley, Zotero, Papers, and othersThe style is either built in or you can download a CSL file that is supported by most references management programs.
BibTeXBibTeX syles are usually part of a LaTeX template. Check the instructions to authors if the publisher offers a LaTeX template for this journal.

Journal articles

Those examples are references to articles in scholarly journals and how they are supposed to appear in your bibliography.

Not all journals organize their published articles in volumes and issues, so these fields are optional. Some electronic journals do not provide a page range, but instead list an article identifier. In a case like this it's safe to use the article identifier instead of the page range.

A journal article with 1 author
Houck O (2003) Tales from a troubled marriage: science and law in environmental policy. Science (New York, N.Y.) 302: 1926–1929.
A journal article with 2 authors
Cyranoski D, Chou I-H (2004) Winds of change blow away the cobwebs on campus. Nature 429: 210–214.
A journal article with 3 authors
Biertümpfel C, Yang W, Suck D (2007) Crystal structure of T4 endonuclease VII resolving a Holliday junction. Nature 449: 616–620.
A journal article with 4 or more authors
Brody DL, Magnoni S, Schwetye KE, Spinner ML, Esparza TJ, Stocchetti N, Zipfel GJ, Holtzman DM (2008) Amyloid-beta dynamics correlate with neurological status in the injured human brain. Science (New York, N.Y.) 321: 1221–1224.

Books and book chapters

Here are examples of references for authored and edited books as well as book chapters.

An authored book
Kitai A (2011) Principles of Solar Cells, LEDs and Diodes: The role of the PN junction. John Wiley & Sons, Ltd, Chichester, UK.
An edited book
Yang X (2012) Incomplete Information System and Rough Set Theory: Models and Attribute Reductions. Yang J (Ed). Springer, Berlin, Heidelberg, XIV, 232 p pp.
A chapter in an edited book
Masera M (2005) Systemic Challenges for Critical Information Infrastructure Protection. In: Kowalik JS, Gorski J, Sachenko A (Eds), Cyberspace Security and Defense: Research Issues: Proceedings of the NATO Advanced Research Workshop on Cyberspace Security and Defense: Research Issues Gdansk, Poland 6–9 September 2004. NATO Science Series II: Mathematics, Physics and Chemistry. Springer Netherlands, Dordrecht, 57–73.

Web sites

Sometimes references to web sites should appear directly in the text rather than in the bibliography. Refer to the Instructions to authors for ZooKeys.

Blog post
Andrews R (2017) Mexican Volcano Puts On Breathtaking Lightning Show. IFLScience. Available from: (October 30, 2018).


This example shows the general structure used for government reports, technical reports, and scientific reports. If you can't locate the report number then it might be better to cite the report as a book. For reports it is usually not individual people that are credited as authors, but a governmental department or agency like "U. S. Food and Drug Administration" or "National Cancer Institute".

Government report
Government Accountability Office (1989) State Department: Problems in the Diplomatic Courier System. U.S. Government Printing Office, Washington, DC

Theses and dissertations

Theses including Ph.D. dissertations, Master's theses or Bachelor theses follow the basic format outlined below.

Doctoral dissertation
Dammann NM (2009) Living in the edge: Community based governance in the aquatic terrestrial zone. Doctoral dissertation. Columbia University

News paper articles

Unlike scholarly journals, news papers do not usually have a volume and issue number. Instead, the full date and page number is required for a correct reference.

New York Times article
Chira S (2017) Mixed Signals on the Runways. New York Times: D1.

In-text citations

References should be cited in the text by name and year in parentheses:

This sentence cites one reference (Houck 2003).
This sentence cites two references (Houck 2003, Cyranoski and Chou 2004).

Here are examples of in-text citations with multiple authors:

  • Two authors: (Cyranoski and Chou 2004)
  • Three or more authors: (Brody et al. 2008)

About the journal

Full journal titleZooKeys
ISSN (print)1313-2989
ISSN (online)1313-2970
ScopeAnimal Science and Zoology
Ecology, Evolution, Behavior and Systematics

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