How to format your references using the ZooKeys citation style

This is a short guide how to format citations and the bibliography in a manuscript for ZooKeys. For a complete guide how to prepare your manuscript refer to the journal's instructions to authors.

Using reference management software

Typically you don't format your citations and bibliography by hand. The easiest way is to use a reference manager:

PaperpileThe citation style is built in and you can choose it in Settings > Citation Style or Paperpile > Citation Style in Google Docs.
EndNoteFind the style here: output styles overview
Mendeley, Zotero, Papers, and othersThe style is either built in or you can download a CSL file that is supported by most references management programs.
BibTeXBibTeX syles are usually part of a LaTeX template. Check the instructions to authors if the publisher offers a LaTeX template for this journal.

Journal articles

Those examples are references to articles in scholarly journals and how they are supposed to appear in your bibliography.

Not all journals organize their published articles in volumes and issues, so these fields are optional. Some electronic journals do not provide a page range, but instead list an article identifier. In a case like this it's safe to use the article identifier instead of the page range.

A journal article with 1 author
Rauchfuss TB (2007) Chemistry. A promising mimic of hydrogenase activity. Science (New York, N.Y.) 316: 553–554.
A journal article with 2 authors
Ausländer S, Fussenegger M (2014) Synthetic biology. Dynamic genome engineering in living cells. Science (New York, N.Y.) 346: 813–814.
A journal article with 3 authors
Dhakal K, Black B, Mohanty S (2014) Introduction of impermeable actin-staining molecules to mammalian cells by optoporation. Scientific reports 4: 6553.
A journal article with 4 or more authors
Blais JM, Kimpe LE, McMahon D, Keatley BE, Mallory ML, Douglas MSV, Smol JP (2005) Arctic seabirds transport marine-derived contaminants. Science (New York, N.Y.) 309: 445.

Books and book chapters

Here are examples of references for authored and edited books as well as book chapters.

An authored book
Chen G, Wang X, Li X (2014) Fundamentals of Complex Networks. John Wiley & Sons Singapore Pte. Ltd, Singapore.
An edited book
Alconada Magliano TM, Chulze SN eds. (2013) Fusarium Head Blight in Latin America. Springer Netherlands, Dordrecht, XX, 304 p. 53 illus., 11 illus. in color pp.
A chapter in an edited book
Lee MS, Chen KW, Ernst E (2010) Supportive Cancer Care with Qigong. In: Cho WCS (Ed), Supportive Cancer Care with Chinese Medicine. Springer Netherlands, Dordrecht, 77–94.

Web sites

Sometimes references to web sites should appear directly in the text rather than in the bibliography. Refer to the Instructions to authors for ZooKeys.

Blog post
Hale T (2016) There Is A Crazy Plan To Use Electricity-Zapping Robots To Hunt Lionfish. IFLScience. Available from: (October 30, 2018).


This example shows the general structure used for government reports, technical reports, and scientific reports. If you can't locate the report number then it might be better to cite the report as a book. For reports it is usually not individual people that are credited as authors, but a governmental department or agency like "U. S. Food and Drug Administration" or "National Cancer Institute".

Government report
Government Accountability Office (1991) Air Force Stock Fund: Hydrazine Sales Consistent with the Commercial Space Launch Act. U.S. Government Printing Office, Washington, DC

Theses and dissertations

Theses including Ph.D. dissertations, Master's theses or Bachelor theses follow the basic format outlined below.

Doctoral dissertation
Tran LN (2013) Low-cost vehicle remote diagnostic system. Doctoral dissertation. California State University, Long Beach

News paper articles

Unlike scholarly journals, news papers do not usually have a volume and issue number. Instead, the full date and page number is required for a correct reference.

New York Times article
Walsh MW (2012) Settlement Money Patched Nassau County Budget Gaps. New York Times: B4.

In-text citations

References should be cited in the text by name and year in parentheses:

This sentence cites one reference (Rauchfuss 2007).
This sentence cites two references (Rauchfuss 2007, Ausländer and Fussenegger 2014).

Here are examples of in-text citations with multiple authors:

  • Two authors: (Ausländer and Fussenegger 2014)
  • Three or more authors: (Blais et al. 2005)

About the journal

Full journal titleZooKeys
ISSN (print)1313-2989
ISSN (online)1313-2970
ScopeAnimal Science and Zoology
Ecology, Evolution, Behavior and Systematics

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