How to format your references using the ZooKeys citation style

This is a short guide how to format citations and the bibliography in a manuscript for ZooKeys. For a complete guide how to prepare your manuscript refer to the journal's instructions to authors.

Using reference management software

Typically you don't format your citations and bibliography by hand. The easiest way is to use a reference manager:

PaperpileThe citation style is built in and you can choose it in Settings > Citation Style or Paperpile > Citation Style in Google Docs.
EndNoteFind the style here: output styles overview
Mendeley, Zotero, Papers, and othersThe style is either built in or you can download a CSL file that is supported by most references management programs.
BibTeXBibTeX syles are usually part of a LaTeX template. Check the instructions to authors if the publisher offers a LaTeX template for this journal.

Journal articles

Those examples are references to articles in scholarly journals and how they are supposed to appear in your bibliography.

Not all journals organize their published articles in volumes and issues, so these fields are optional. Some electronic journals do not provide a page range, but instead list an article identifier. In a case like this it's safe to use the article identifier instead of the page range.

A journal article with 1 author
Sherwood S (2002) A microphysical connection among biomass burning, cumulus clouds, and stratospheric moisture. Science (New York, N.Y.) 295: 1272–1275.
A journal article with 2 authors
Kaspi Y, Schneider T (2011) Winter cold of eastern continental boundaries induced by warm ocean waters. Nature 471: 621–624.
A journal article with 3 authors
Sergio F, Newton I, Marchesi L (2005) Conservation: top predators and biodiversity. Nature 436: 192.
A journal article with 4 or more authors
Wooding S, Bufe B, Grassi C, Howard MT, Stone AC, Vazquez M, Dunn DM, Meyerhof W, Weiss RB, Bamshad MJ (2006) Independent evolution of bitter-taste sensitivity in humans and chimpanzees. Nature 440: 930–934.

Books and book chapters

Here are examples of references for authored and edited books as well as book chapters.

An authored book
Howe TR (2011) Marriages & Families in the 21st Century. Wiley-Blackwell, Oxford, UK.
An edited book
Lhoneux E de (2014) The European Stability Mechanism Before the Court of Justice of the European Union: Comments on the Pringle Case. Vassilopoulos CA (Ed). Springer International Publishing, Cham, VIII, 74 p pp.
A chapter in an edited book
Ono T, Ambe D, Kaeriyama H, Shigenobu Y, Fujimoto K, Sogame K, Nishiura N, Fujikawa T, Morita T, Watanabe T (2015) Radiocesium Concentrations in the Organic Fraction of Sea Sediments. In: Nakata K, Sugisaki H (Eds), Impacts of the Fukushima Nuclear Accident on Fish and Fishing Grounds. Springer Japan, Tokyo, 67–75.

Web sites

Sometimes references to web sites should appear directly in the text rather than in the bibliography. Refer to the Instructions to authors for ZooKeys.

Blog post
Andrew E (2015) Awesome Video Shows A Soyuz Launching And Docking To The International Space Station. IFLScience. Available from: (October 30, 2018).


This example shows the general structure used for government reports, technical reports, and scientific reports. If you can't locate the report number then it might be better to cite the report as a book. For reports it is usually not individual people that are credited as authors, but a governmental department or agency like "U. S. Food and Drug Administration" or "National Cancer Institute".

Government report
Government Accountability Office (2007) Weather Forecasting: National Weather Service’s Operations Prototype Needs More Rigorous Planning. U.S. Government Printing Office, Washington, DC

Theses and dissertations

Theses including Ph.D. dissertations, Master's theses or Bachelor theses follow the basic format outlined below.

Doctoral dissertation
Karmacharya B (2015) Population Dynamics of Northern Cardinal and Carolina Wren in an Urban Forest Fragment: Safe Refuge or Ecological Trap? Doctoral dissertation. University of Louisiana

News paper articles

Unlike scholarly journals, news papers do not usually have a volume and issue number. Instead, the full date and page number is required for a correct reference.

New York Times article
Koblin J (2017) New Life Emerges From the “Big Bang.” New York Times: AR114.

In-text citations

References should be cited in the text by name and year in parentheses:

This sentence cites one reference (Sherwood 2002).
This sentence cites two references (Sherwood 2002, Kaspi and Schneider 2011).

Here are examples of in-text citations with multiple authors:

  • Two authors: (Kaspi and Schneider 2011)
  • Three or more authors: (Wooding et al. 2006)

About the journal

Full journal titleZooKeys
ISSN (print)1313-2989
ISSN (online)1313-2970
ScopeAnimal Science and Zoology
Ecology, Evolution, Behavior and Systematics

Other styles