How to format your references using the ZooKeys citation style

This is a short guide how to format citations and the bibliography in a manuscript for ZooKeys. For a complete guide how to prepare your manuscript refer to the journal's instructions to authors.

Using reference management software

Typically you don't format your citations and bibliography by hand. The easiest way is to use a reference manager:

PaperpileThe citation style is built in and you can choose it in Settings > Citation Style or Paperpile > Citation Style in Google Docs.
EndNoteFind the style here: output styles overview
Mendeley, Zotero, Papers, and othersThe style is either built in or you can download a CSL file that is supported by most references management programs.
BibTeXBibTeX syles are usually part of a LaTeX template. Check the instructions to authors if the publisher offers a LaTeX template for this journal.

Journal articles

Those examples are references to articles in scholarly journals and how they are supposed to appear in your bibliography.

Not all journals organize their published articles in volumes and issues, so these fields are optional. Some electronic journals do not provide a page range, but instead list an article identifier. In a case like this it's safe to use the article identifier instead of the page range.

A journal article with 1 author
Pyne SJ (2001) Essays on science and society. The fires this time, and next. Science (New York, N.Y.) 294: 1005–1006.
A journal article with 2 authors
Campbell E, Salathé M (2013) Complex social contagion makes networks more vulnerable to disease outbreaks. Scientific reports 3: 1905.
A journal article with 3 authors
Giles S, Friedman M, Brazeau MD (2015) Osteichthyan-like cranial conditions in an Early Devonian stem gnathostome. Nature 520: 82–85.
A journal article with 4 or more authors
Zhang X, Fu W, Palivan CG, Meier W (2013) Natural channel protein inserts and functions in a completely artificial, solid-supported bilayer membrane. Scientific reports 3: 2196.

Books and book chapters

Here are examples of references for authored and edited books as well as book chapters.

An authored book
Schepers U (2004) RNA Interference in Practice. Wiley-VCH Verlag GmbH & Co. KGaA, Weinheim, FRG.
An edited book
Dobrovie-Sorin C (2012) 85 Redefining Indefinites. Beyssade C (Ed). Springer Netherlands, Dordrecht, XVI, 268 p pp.
A chapter in an edited book
Francesconi F, Dalpiaz F, Mylopoulos J (2013) TBIM: A Language for Modeling and Reasoning about Business Plans. In: Ng W, Storey VC, Trujillo JC (Eds), Conceptual Modeling: 32th International Conference, ER 2013, Hong-Kong, China, November 11-13, 2013. Proceedings. Lecture Notes in Computer Science. Springer, Berlin, Heidelberg, 33–46.

Web sites

Sometimes references to web sites should appear directly in the text rather than in the bibliography. Refer to the Instructions to authors for ZooKeys.

Blog post
Davis J (2015) How Do Deadly Mites Trick Honeybees? IFLScience. Available from: (October 30, 2018).


This example shows the general structure used for government reports, technical reports, and scientific reports. If you can't locate the report number then it might be better to cite the report as a book. For reports it is usually not individual people that are credited as authors, but a governmental department or agency like "U. S. Food and Drug Administration" or "National Cancer Institute".

Government report
Government Accountability Office (2016) Special Education: State and Local-Imposed Requirements Complicate Federal Efforts to Reduce Administrative Burden. U.S. Government Printing Office, Washington, DC

Theses and dissertations

Theses including Ph.D. dissertations, Master's theses or Bachelor theses follow the basic format outlined below.

Doctoral dissertation
Moulder JC (2010) Comparative analgesic effects of prepulse inhibition and reduced voltage waveforms in defibrillation. Doctoral dissertation. George Washington University

News paper articles

Unlike scholarly journals, news papers do not usually have a volume and issue number. Instead, the full date and page number is required for a correct reference.

New York Times article
Wasik JF (2016) Helping Hand for Start-Ups From Those Who’ve Been There Before. New York Times: B2.

In-text citations

References should be cited in the text by name and year in parentheses:

This sentence cites one reference (Pyne 2001).
This sentence cites two references (Pyne 2001, Campbell and Salathé 2013).

Here are examples of in-text citations with multiple authors:

  • Two authors: (Campbell and Salathé 2013)
  • Three or more authors: (Zhang et al. 2013)

About the journal

Full journal titleZooKeys
ISSN (print)1313-2989
ISSN (online)1313-2970
ScopeAnimal Science and Zoology
Ecology, Evolution, Behavior and Systematics

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