How to format your references using the ZooKeys citation style

This is a short guide how to format citations and the bibliography in a manuscript for ZooKeys. For a complete guide how to prepare your manuscript refer to the journal's instructions to authors.

Using reference management software

Typically you don't format your citations and bibliography by hand. The easiest way is to use a reference manager:

PaperpileThe citation style is built in and you can choose it in Settings > Citation Style or Paperpile > Citation Style in Google Docs.
EndNoteFind the style here: output styles overview
Mendeley, Zotero, Papers, and othersThe style is either built in or you can download a CSL file that is supported by most references management programs.
BibTeXBibTeX syles are usually part of a LaTeX template. Check the instructions to authors if the publisher offers a LaTeX template for this journal.

Journal articles

Those examples are references to articles in scholarly journals and how they are supposed to appear in your bibliography.

Not all journals organize their published articles in volumes and issues, so these fields are optional. Some electronic journals do not provide a page range, but instead list an article identifier. In a case like this it's safe to use the article identifier instead of the page range.

A journal article with 1 author
Shastri N (2006) Cell biology. Peptides, scrambled and stitched. Science (New York, N.Y.) 313: 1398–1399.
A journal article with 2 authors
Gal-Yam A, Leonard DC (2009) A massive hypergiant star as the progenitor of the supernova SN 2005gl. Nature 458: 865–867.
A journal article with 3 authors
Thammavongsa V, Missiakas DM, Schneewind O (2013) Staphylococcus aureus degrades neutrophil extracellular traps to promote immune cell death. Science (New York, N.Y.) 342: 863–866.
A journal article with 4 or more authors
Pennycook TJ, Jones L, Pettersson H, Coelho J, Canavan M, Mendoza-Sanchez B, Nicolosi V, Nellist PD (2014) Atomic scale dynamics of a solid state chemical reaction directly determined by annular dark-field electron microscopy. Scientific reports 4: 7555.

Books and book chapters

Here are examples of references for authored and edited books as well as book chapters.

An authored book
Padilla E (2015) Substation Automation Systems. John Wiley & Sons, Ltd, Chichester, UK.
An edited book
Moehrle MG, Isenmann R, Phaal R eds. (2013) Technology Roadmapping for Strategy and Innovation: Charting the Route to Success. Springer, Berlin, Heidelberg, VIII, 285 p pp.
A chapter in an edited book
Amorim F, Moseley PL (2010) Heat Shock Protein and Inflammation. In: Asea AAA, Pedersen BK (Eds), Heat Shock Proteins and Whole Body Physiology. Heat Shock Proteins. Springer Netherlands, Dordrecht, 57–83.

Web sites

Sometimes references to web sites should appear directly in the text rather than in the bibliography. Refer to the Instructions to authors for ZooKeys.

Blog post
Andrew E (2015) Busting Myths: A Practical Guide To Countering Science Denial. IFLScience. Available from: (October 30, 2018).


This example shows the general structure used for government reports, technical reports, and scientific reports. If you can't locate the report number then it might be better to cite the report as a book. For reports it is usually not individual people that are credited as authors, but a governmental department or agency like "U. S. Food and Drug Administration" or "National Cancer Institute".

Government report
Government Accountability Office (1991) Railroad Safety: DOD Can Improve the Safety of On-Base Track and Equipment. U.S. Government Printing Office, Washington, DC

Theses and dissertations

Theses including Ph.D. dissertations, Master's theses or Bachelor theses follow the basic format outlined below.

Doctoral dissertation
Lee JY (2008) Socio-Spatial Exclusion Based on Human Activities and Social Networks in Space-Time: A Case Study of Koreans in Columbus, Ohio. Doctoral dissertation. Ohio State University

News paper articles

Unlike scholarly journals, news papers do not usually have a volume and issue number. Instead, the full date and page number is required for a correct reference.

New York Times article
Vecsey G (2014) Jerry Coleman, 89, Yankee Infielder, Fighter Pilot and Voice of the Padres. New York Times: A15.

In-text citations

References should be cited in the text by name and year in parentheses:

This sentence cites one reference (Shastri 2006).
This sentence cites two references (Shastri 2006, Gal-Yam and Leonard 2009).

Here are examples of in-text citations with multiple authors:

  • Two authors: (Gal-Yam and Leonard 2009)
  • Three or more authors: (Pennycook et al. 2014)

About the journal

Full journal titleZooKeys
ISSN (print)1313-2989
ISSN (online)1313-2970
ScopeAnimal Science and Zoology
Ecology, Evolution, Behavior and Systematics

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